''Polystichum munitum'', the western swordfern, is an

evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...

perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...

native to western

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

, where it is one of the most abundant ferns in forested areas. It occurs along the Pacific coast from southeastern

Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...

to southern

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

, and also inland east to southeastern

British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...

, northern

Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyom ...

and western

Montana Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...

, with disjunctive populations in northern

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

; the

Black Hills The Black Hills ( lkt, Ȟe Sápa; chy, Moʼȯhta-voʼhonáaeva; hid, awaxaawi shiibisha) is an isolated mountain range rising from the Great Plains of North America in western South Dakota and extending into Wyoming, United States. Black Elk P ...

South Dakota South Dakota (; Sioux language, Sioux: , ) is a U.S. state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Lakota people, Lakota and Dakota peo ...

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

; and

Guadalupe Island Guadalupe Island ( es, Isla Guadalupe, link=no) is a volcanic island located off the western coast of Mexico's Baja California Peninsula and about southwest of the city of Ensenada in the state of Baja California, in the Pacific Ocean. The ...

off of

Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

. Western swordfern is known to have locally

naturalized Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...

in parts of

Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...

and

Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...

Description

The dark green

frond A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the lar ...

s of this fern grow tall, in a tight clump spreading out radially from a round base. They are single-

pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...

, with the pinnae alternating on the stalk. Each pinna is long, with a small upward-pointing lobe (a sword hilt, hence the name) at the base, and the edges are serrated with bristly tips. Individual fronds live for 1.5 to 2.5 years and remain attached to the

rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...

after withering. The round sori occupy two rows on either side of the midrib of each pinna and are covered by a centrally-attached, umbrella-like indusium with fringed edges. They produce light yellow spores. In deep shade, fronds spread nearly horizontally but with increasing sun exposure grow more upright. Hitchcock, C.L. and Cronquist, A. 2018. Flora of the Pacific Northwest, 2nd Edition, p. 56. University of Washington Press, Seattle.

Habitat and cultivation

The preferred habitat of this fern is the understory of moist

conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...

ous woodlands at low elevations. It grows best in well-drained acidic

soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...

of rich

humus In classical soil science, humus is the dark organic matter in soil that is formed by the decomposition of plant and animal matter. It is a kind of soil organic matter. It is rich in nutrients and retains moisture in the soil. Humus is the Lati ...

and small stones. It is very resilient and survives occasional droughts, but flourishes only with consistent moisture and it prefers cool weather. In cultivation, it also responds well to regular, light fertilization. While this fern is a favored horticultural subject in western

, it has proved difficult or impossible to cultivate satisfactorily in the eastern part of the continent. In the

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...

''Polystichum munitum'' has gained the

Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nort ...

Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...

Utility

The Coast Salish people of B.C. and Washington state use this plant as a pain reliever. When applied directly to the area where pain and inflammation occur, according to Della Rice Sylvester, an elder and medicine woman of the Cowichan tribe, the sword fern "takes the pain away!". This traditional use has spread among the hiking communities and youth scouting organizations of the region, where it is a common piece of hiker's lore that a rash from a stinging nettle can be counteracted by rubbing the spores on the underside of sword fern on the area. In spring, when other food is unavailable,

Quileute The Quileute , are a Native American people in western Washington state in the United States, currently numbering approximately 2,000. They are a federally recognized tribe: the ''Quileute Tribe of the Quileute Reservation''. The Quileute peo ...

Makah The Makah (; Klallam: ''màq̓áʔa'')Renker, Ann M., and Gunther, Erna (1990). "Makah". In "Northwest Coast", ed. Wayne Suttles. Vol. 7 of ''Handbook of North American Indians'', ed. William C. Sturtevant. Washington, D.C.: Smithsonian Institut ...

Klallam Klallam (also Clallam, although the spelling with "K" is preferred in all four modern Klallam communities) refers to four related indigenous Native American/First Nations communities from the Pacific Northwest of North America. The Klallam cult ...

, Squamish,

Sechelt Sechelt (, Shishalh language chat'lich) is a district municipality located on the lower Sunshine Coast of British Columbia. Approximately 50 km northwest of Vancouver, it is accessible from mainland British Columbia by a 40-minute ferry tr ...

, Haida, and other Native American/

First Nations First Nations or first peoples may refer to: * Indigenous peoples, for ethnic groups who are the earliest known inhabitants of an area. Indigenous groups *First Nations is commonly used to describe some Indigenous groups including: **First Natio ...

peoples roasted, peeled, and ate the

s. The fronds were used to line fire pits and food drying racks and as filling for mattresses. The plant is also cultivated for its ornamental foliage, which

florist Floristry is the production, commerce, and trade in flowers. It encompasses flower care and handling, floral design and arrangement, merchandising, production, display and flower delivery. Wholesale florists sell bulk flowers and related sup ...

s include in vases.

References

External links

USDA Plants Profile: ''Polystichum munitum''

Jepson eFlora Treatment of ''Polystichum munitum''

Calflora Database: ''Polystichum munitum'' (western sword fern)

Flora of North America: ''Polystichum munitum''

{{Taxonbar, from=Q2273822 munitum Ferns of California Flora of the West Coast of the United States Flora of the Northwestern United States Flora of Alaska Flora of Baja California Flora of Mexican Pacific Islands Flora of California Flora of British Columbia Flora of Montana Flora of Oregon Flora of South Dakota Flora of the Cascade Range Flora of the Great Basin Flora of the Klamath Mountains Flora of the Sierra Nevada (United States) Natural history of the California chaparral and woodlands Natural history of the California Coast Ranges Natural history of the Channel Islands of California Natural history of the San Francisco Bay Area Taxa named by Carl Borivoj Presl Garden plants of North America Flora without expected TNC conservation status